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Live BBC News coverage of the European election results
Nigel Farage has said his "dream has become a reality" and UKIP is now the "third force" in British politics after it topped the European polls.
With only Scotland left to declare, UKIP has 27.5% of the vote and 23 MEPs.
Labour, on 25%, is narrowly beating the Tories into second place while the Lib Dems lost all but one of their seats and came behind the Greens.
David Cameron said the public was "disillusioned" with the EU and their message was "received and understood".
The full Scottish result will be known at noon on Monday, as the Western Isles does not count votes on a Sunday.
The BBC is predicting that the result in Scotland will be SNP on two seats, Labour on two seats and the Conservatives and UKIP with one seat each - meaning Mr Farage's anti-EU party is expected to win at least one seat in every part of Great Britain.
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Nigel Farage: "The people's army of UKIP have spoken tonight"
It will be the first time a national election has not been won by the Conservatives or Labour in 100 years - and the first time a party with no MPs at Westminster has achieved such a result.
Counting in Northern Ireland begins later on Monday.
- UKIP topped the poll in six of the 10 regions to declare, with its strongest performance coming in the East Midlands, where its vote was up 16.5% to 33%
- Labour topped the poll in Wales, the North-West of England, the North-East of England and London where it increased its share of the vote by 15% to 36.7%
- Ed Miliband's party has 18 MEPs so far, an increase of seven on 2009, which was a record low point for the party
- UKIP came third in London with 16.9% of the vote
- The Conservatives have so far secured 24% of the vote nationally and lost seven seats
- The Lib Dems have lost nine MEPs but avoided a humiliating wipe-out by winning a single seat in the South-East of England
- The Green Party has got three MEPs - one more than it achieved in 2009
- Coverage is on the BBC News Channel and BBC World. You can follow all the latest news, reaction and results on bbc.co.uk/vote2014
In his victory speech at the South East of England count, UKIP leader Mr Farage said: "The people's army of UKIP have spoken tonight and have delivered just about the most extraordinary result in British politics for 100 years."
End QuoteIt is over 100 years since a national election has been won by a party other than the Conservatives and Labour"
He said the three main parties in Britain had led the country into the Common Market but had "twisted and turned" over an in/out referendum on EU membership.
"The penny's really dropped that as members of this union we cannot run our own country and crucially, we cannot control our own borders," said the UKIP leader.
He later told the BBC that he was "over the moon" about his party's performance and he was more confident than ever that it would make a breakthrough at Westminster next year.
The result had "serious consequences" for the three main party leaders and he believed Nick Clegg's position as Lib Dem leader was now "untenable".
Mr Clegg faced calls to stand down ahead of Sunday night's results - something firmly rejected by senior party figures.
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Chief Secretary to the Treasury Danny Alexander (R) rejects calls from federal executive member Martin Tod (L) for a new party leader
A member of the party's federal executive, Martin Tod, reiterated his call for Mr Clegg to quit because voters were "not prepared" to listen to him.
He said any new leader should "prove themselves in a democratic party election".
Mr Tod, a councillor in Winchester, branded his party's results in the European elections a "disaster" and the party's response "complacent".
But Treasury Chief Secretary Danny Alexander, who has been touted as a future leader himself, insisted Mr Clegg was "by far the best spokesman" for the Lib Dems and said "plunging the party into a period of introspection" was not the right response to the results.
Conservative party chairman Grant Shapps said the results were acting as "a command for Britain to get a better deal" in Europe - but he rejected calls by Tory grandee David Davis to bring forward the in/out EU referendum to 2016, saying negotiations on this could not be rushed.
The Tory leadership faced further calls for an electoral pact with UKIP to avoid a split in the right of British politics at next year's general election.
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Foreign Secretary William Hague: "We can fix the relationship between Britain and Europe"
Daniel Hannan, who was returned as a Tory MEP in the South East region, said: "It would be very sad if the two right-of-centre Eurosceptic parties at the general election were not able to find some way, at least in marginal seats, of reaching an accommodation so that anti-referendum candidates don't get in with a minority of votes."
Grant Shapps has repeatedly ruled out any formal pact or local deals with UKIP.
Labour was looking at one stage as if it might be beaten into third place by the Tories - a potentially disastrous result for Ed Miliband as he seeks to show he can win next year's general election.
But the party was rescued by another strong showing in London - and it took heart from local election results in battleground seats, which party spokesmen suggested were a better guide to general election performance.
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Natalie Bennett: "What people are saying... is that [they're] not happy with politics as it is now"
Labour deputy leader Harriet Harman said UKIP had tapped into people's disaffection with politics.
She told BBC people have said they are voting UKIP, even if they are a Labour supporter, because, they said: "I think you need a shake-up."
BNP leader Nick Griffin lost his seat and saw his party's vote collapse by 6% in the North-West of England.
The BNP has seen a similar slump in support across the UK and seems certain to have no representation in the next European Parliament.
Across the channel, the far right anti-EU National Front has topped the poll in France, according to exit polls.
Anti-EU parties from the left and right are expected to gain significant numbers of MEPs across all 28 member states in the wake of the eurozone crisis and severe financial squeeze.
But pro-EU parties are still expected to hold the majority in parliament.
Turnout across the EU is up slightly at 43.1%, according to estimates. Turnout in the UK was 33.8%, down slightly on last time.
In the European elections five years ago, the Conservatives got 27.7% of the total vote, ahead of UKIP on 16.5%, Labour on 15.7%, the Lib Dems on 13.7%, the Green Party on 8.6% and the BNP on 6.2%.
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